The Lemp sisters, Emma (Gale Page), Thea (Lola Lane), Kay (Rosemary Lane), and Ann (Priscilla Lane) are prodigies in a musical family headed by their father, Adam (Claude Rains). The Lemps also run a boarding house, and among the tenants is Felix Deitz (Jeffrey Lynn), a young composer whom the four daughters want to attract.
Emma, the oldest daughter, is the object of affection of a neighbor, Ernest (Dick Foran), but she rebuffs his attentions. Thea, a pianist and the second eldest, is courted by wealthy Ben Crowley (Frank McHugh), another neighbor, but she is not sure she loves him. Kay, the third daughter, is a talented singer and has a chance at a music school scholarship but doesn't want to leave home. The youngest daughter is Ann, a violinist. Mickey (John Garfield), an orchestral arranger and friend of Felix, falls for Ann, but Felix also has had his eyes on her and proposes marriage.
An hour before Ann and Felix are to marry, Mickey tells Ann that Emma is in love with Felix. Ann then observes Emma's distress after parting from Felix for the ceremony, and decides to step out of their way. She leaves and sends a telegram the wedding participants get, telling them she has eloped with Mickey. Ernest is prominent in calming people.
Four months later, Ann and Mickey are living a hard life in New York, professing love for each other but penniless and unhappy. Mickey is invited to form a band and go to South America with some fellow-musicians, but cannot afford passage, and Ann does not want to go.
The family meets at the Lemps' house for Christmas, except Kay, who is singing on the radio that night. Emma has not gotten together with Felix, but is now engaged to Ernest; she tells Ann that she had thought she was in love with Felix and would have kept on thinking so and spoiled her life if Ann had married him, but she was awakened to Ernest's qualities when he took charge at the abortive wedding. Felix is alone and unhappy, though he has a good conducting job awaiting him in Seattle. Mickey observes all this, and feels like an outsider and a failure. When he and Ben go to see Felix off at the train station in a snowstorm, he drives Ben's car and drops Ben off for an errand; as the train pulls out, Felix presses some money on him and tells him to use it for Ann, and Mickey, realizing that Felix and Ann really belong together, speeds through the snow until he crashes the car. The family comes to the hospital, and he dies with Ann at his bedside.
A while later, all four daughters are home making music with their loved ones when Felix returns, and he and Ann renew their love.
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