In Siberia (or possibly Manchuria or even Szechuan province), 1906, anthropologist Alexander Saxton (Lee) finds a frozen creature which he believes to be the Missing Link. He transports the being back to western Europe by trans-Siberian train. Aboard the train are the usual assorted types:- a Polish countess (Silvia Tortosa); her husband (Jorge Rigaud); a mad monk seemingly modelled on Rasputin (Alberto De Mendoza); and a fellow scientist and long-time rival of Saxton's, the charismatic Dr Wells (Cushing). Midway through the journey, as the train passes through a desolate snowscape many miles from civilisation, the creature thaws out and begins killing the passengers. Wells carries out an autopsy on the corpses and realises that they are not up against the Missing Link, but a weird alien organism which inhabits the bodies of its victims and steals their memories. As the finger of suspicion falls upon each character, the scientists try to figure out which passenger is "carrying" the shape-shifting monster, while trying to keep everyone safe and alive.
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